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| | Journal of Aquariculture & Aquatic Sciences Article |
 | | Chemical signals play a part in initiating nest building activity in anabantid males (Rossi, 1969) while pre-embrace physical contact (in the form of biting, butting and "beating" with the pelvic fins) contributes to successful mating (Forselius, 1957; Hall and Miller, 1968). |
 | | Successful mating in anabantids, therefore, relies on complex interplay between visual, chemical (pheromonal), physical and mechanical (touch) signals. |
 | | The female's eyes had, presumably, been lost through injury, such as might be suffered as the result of an attack by a larger, predatory fish in the close confines of an aquarium. |
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