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 | | Fisher showed, by using a diffusion model, that after a gene was established in a deme there would be a wave of advance for which V= (1/2)s(s)1/2, where V is the velocity of the wave, and s is the selective advantage of the allele. |
 | | He provides evidence, from the spread of Carpentarian characteristics in Australia, that genetic diffusion is indeed as would be predicted from a model where tribal boundaries are an obstacle to gene flow, and the boundaries are further apart in areas of low rainfall. |
 | | Thus, the rate of diffusion of genes would be limited by the boundaries of the tribal groups, and the short distances from one boundary to the other would again limit the gene diffusion. |
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