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| | ..::treeBASE::.. (Site not responding. Last check: 2007-10-10) |
 | | Important points of uncertainty or contention include, apart from ranking problems, monophyly of Papilionoidea plus Hesperioidea, their relationship with other Lepidoptera in general and the Hedyloidea in particular, the question of the sister group of the Pieridae (either Papilionidae, or Lycaenidae + Nymphalidae), and the division of families into subfamilies. |
 | | Traditional groupings are discussed and compared with the results of a cladistic analysis using 103 characters and 74 species (59 butterflies and 15 moths). |
 | | Nevertheless, of the moths used, Macrosoma (Hedylidae, Hedyloidea) and Urania (Uraniidae, Geometroidea) appear to be the closest relatives of the butterflies. |
| www.phylo.org /treebase/view/view_study.php?studyID=S471 (338 words) |
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